Maximum clade credibility chronogram estimated by BEAST. The rhombus (◇) and star (☆) signs denote calibration points for molecular dating. Dashed clades indicate PP < 0.75 and thicken clades indicate PP ≥ 0.95. Node heights indicate mean ages, with 95% highest posterior density (HPD) date ranges shown by the node bars. Clades with PP ≥ 0.75 are numbered and their clade support and divergent ages mean (95% HPD) are shown in Table 2.